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I think what is being referred to here is how light levels affect mood. It is known that sunlight levels can greatly affect mood, sleep patterns, etc. In places in the world where little sunlight is received for parts of the year, depression can be a problem if not looked into.
In WoW, the Draenei starting area, Azuremyst Isle, and the area Darkshore**, are both really gray, and I personally think they they are the most "depressing" areas in WoW, far more so than any fiery/doomy high-level area, because they are so desolate, but I also think because they are so sparsely populated.
The point I am making is that even if an area is gloomy, if it is fairly well populated with other players to talk and LOL with, and you kept are busy questing and fighting, you may be less likely to fall into the dark moods the areas evoke.
(**Both I think are patterned on the beaches of the Pacific Northwest USA, like Washington state, which is pretty bleak.)
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